@booklet {121, title = {Towards a monophyletic Omphalodes{\textemdash}or an expansion of North American Mimophytum}, howpublished = {PhytotaxaPhytotaxa}, volume = {288}, number = {2}, year = {2016}, pages = {131{\textendash}144}, abstract = {

Molecular studies have recently led to major changes to the systematics of Boraginaceae and changed our perception of fruit morphology, previously used as the most important character in Boraginaceae classification. Navel-shaped nutlets as the key character for Omphalodes turned out to be highly homoplasious, thus the genus was revealed to be polyphyletic. After the recent reclassifications, the genus Omphalodes is still left paraphyletic, since the remaining taxa correspond to two different lineages: a clade of essentially Mediterranean perennials including the type species (O. verna) and a clade with the annual to subperennial North American taxa including the genus Mimophytum . We here advocate transferring all taxa of the North American clade into the genus Mimophytum, subdividing the genus Omphalodes into morphologically coherent and monophyletic entities. Mimophytum was initially removed from Omphalodes based on its glochidiate nutlets, but this character state does not define a monophyletic entity. In overall morphology, the redefined genus Mimophytum is highly homogeneous and easily differentiated from the other clades: it has a lax growth habit, and leaves are long-petiolate, abruptly contracted at the base into a more or less cordate base, and with a fine pubescence of erect trichomes. After the proposed rearrangements, only O. erecta remains as a problematic species whose systematic placement is still to be found out.

}, keywords = {Eudicots, Boraginaceae, navel seeds, Sierra Madre Oriental}, doi = {10.11646/phytotaxa.288.2.3}, author = {Holstein, Norbert and Chac{\'o}n, Juliana and Otero, Ana and Jim{\'e}nez-Mej{\'\i}as, Pedro and M. Weigend} } @article {119, title = {Lectotypifications and nomenclatural notes in the Boraginaceae s.str. for the Flora of Ecuador}, journal = {Phytotaxa}, volume = {278}, number = {1}, year = {2016}, month = {05/10/2016}, pages = {73}, chapter = {69}, abstract = {

During the preparation of a taxonomic treatment concerning Boraginaceae s.str. for the Flora of Ecuador, we found that there are eleven species currently known in that area. We herewith designate lectotypes for some of the names concerned: Cynoglossum revolutum and Cynoglossum mexicanum (both Hackelia revoluta), Amsinckia hispida and Eritrichium mandonii (both Amsinckia calycina), and Anchusa linifolia and Lithospermum alpinum (both Plagiobothrys linifolius). The complex history of the type material of various synonyms of Plagiobothrys linifolius based on material by Bonpland and Humboldt is briefly discussed.

}, keywords = {Bonpland, Eudicots, Humboldt, Kunth, Ruiz L{\'o}pez \& Pav{\'o}n, South America, typification, Willdenow}, doi = {10.11646/phytotaxa.278.1.9}, url = {http://biotaxa.org/Phytotaxa/article/view/phytotaxa.278.1.9}, author = {Holstein, Norbert and M. Weigend} } @booklet {116, title = {No longer shipwrecked{\textemdash}Selkirkia (Boraginaceae) back on the mainland with generic rearrangements in South American {\textquotedblleft}Omphalodes{\textquotedblright} based on molecular data}, howpublished = {PhytotaxaPhytotaxa}, volume = {270}, number = {4}, year = {2016}, month = {24 Aug 2016}, pages = {231{\textendash}251}, publisher = {Magnolia Press}, edition = {24 Aug 2016}, chapter = {231}, abstract = {

The genus Omphalodes (Boraginaceae) has recently been shown to be polyphyletic. Two distantly related lineages have already been segregated into the genera Memoremea (Central Europe) and Nihon (East Asia), respectively. We expanded the taxon sampling in the Omphalodeae and confirm that the genus is still paraphyletic to the two monotypic genera Selkirkia from the Juan Fern\ández Islands off the coast of Chile and Myosotidium from Chatham Island off the coast of New Zealand, plus two South American species currently assigned to Cynoglossum, and one species recently segregated from the latter genus as Mapuchea. Four clades are retrieved in a narrowly delimited Omphalodes group: 1) Iberodes, the annual southwestern European species of Omphalodes s.l. that have been recently segregated into this genus, 2) Omphalodes s.str., perennial western Eurasian species (including the type species of the genus), 3) the North American species of Omphalodes, and 4) the southern hemispheric Myosotidium as sister to a monophyletic group with Mapuchea plus the two other South American species of \“Cynoglossum\” and the island shrub Selkirkia berteroi. We argue that the taxa of this latter clade are best placed into an expanded genus Selkirkia. Selkirkia then represents a morphologically coherent entity with glochidiate nutlets. Its considerable difference in vegetative morphology to Myosotidium is easily explained by the highly divergent habitats the respective plants occupy. Lectotypifications, illustrations, and descriptions are provided for Myosotidium and the species of the expanded genus Selkirkia.

}, keywords = {island woodiness, ITS, molecular phylogeny, Taxonomy, trnL{\textendash}trnL{\textendash}F}, author = {Holstein, Norbert and Chac{\'o}n, Juliana and Hilger, Hartmut H. and M. Weigend} }