01409nas a2200145 4500008004100000245004700041210004700088300001400135490000800149520101200157100001301169700001501182700001901197856004701216 2002 eng d00aTransfer cells in the seeds of Boraginales0 aTransfer cells in the seeds of Boraginales a155–1640 v1403 a

The presence of transfer cells (TCs) in the seeds of Boraginales (Boraginaceae s.s., Hydrophyllaceae, Heliotropiaceae, Ehretiaceae, Cordiaceae and Lennoaceae) has been reported but has not hitherto been studied systematically. This study, surveying the seed anatomy of 50 species of Boraginales, demonstrates that in Heliotropiaceae, Cordiaceae, Ehretiaceae and Lennoaceae, TCs are found in an uninterrupted course from the placenta via the funicle to the seed coat. These families are characterized by indehiscent fruits with a protective endocarp. The TCs may act as a sponge, thus promoting rapid germination when sufficient water is available. In Hydrophyllaceae, which have capsular fruits, TCs are often found but have a different structure and are restricted to the seed coat. Boraginaceae s.s. on the other hand are characterized by a complete absence of TCs in their seeds. The presence of TCs in seeds is considered as a synapomorphic trait and appears to be phylogenetically informative.

1 aDiane, N1 aHilger, HH1 aGottschling, M uhttps://boraginales.myspecies.info/node/8000466nas a2200121 4500008004100000245013500041210007000176300001400246490000700260100001500267700001500282856004700297 1982 eng d00aUntersuchungen zur Ausbildung der Klausen und ihrer Oberflächenskulpturen bei Paracaryum intermedium (Boraginaceae) aus Jordanien0 aUntersuchungen zur Ausbildung der Klausen und ihrer Oberflächens a205–2200 v571 aHilger, HH1 aRichter, U uhttps://boraginales.myspecies.info/node/5601219nas a2200145 4500008004100000245020600041210007000247300001400317490000800331520065200339100001500991700000501006700001501011856004701026 1993 eng d00aEnergiedispersive Röntgenmikroanalyse (EDX) von Boraginaceae subfam. Boraginoideae - Klausenoberflachen. Sind Silicium- und Calcium - Einlagerungen in die Fruchtwand systematisch verwertbare Merkmale?0 aEnergiedispersive Röntgenmikroanalyse EDX von Boraginaceae subfa a387–3980 v1883 a

Energy dispersive X-ray microanalysis (EDX) was used to investigate surfaces of Boraginoideae nutlets with regard to their content and ratios of silicon (Si) and calcium (Ca), respectively. Deposition takes place very short prior to full maturity. Potassium (K) is a third always detectible element.
It is not possible to associate species to a tribe only with respect to different element contents. Morphology and sculpturing of nutlets play a more important role. The Lithospermeae which exhibit the most elaborate pericarp anatomy have a hight amount of Ca deposition in the lumina of the cells of the outermost mesocarp layer.

1 aHilger, HH1 a1 aHofmann, M uhttps://boraginales.myspecies.info/node/5702073nas a2200133 4500008004100000245009600041210006900137300001200206490000800218520163800226100001501864700001301879856004701892 2003 eng d00aA systematic analysis of Heliotropiaceae (Boraginales) based on trnL and ITS1 sequence data0 asystematic analysis of Heliotropiaceae Boraginales based on trnL a19–510 v1253 a

The infrafamilial relationships of Heliotropiaceae (Boraginaceae subfam. Heliotropioideae according to traditional systems) are reevaluated using molecular data of nuclear ITS1 (86 species) and plastidal trn LUAA intron (66 species) sequences. The results obtained from our investigations show that traditional generic limits warrant adjustment. Heliotropiaceae fall into two large clades. The first clade includes, in basal position, the genus Ixorhea . The genus Myriopus (formerly Tournefortia sect. Cyphocyema) is sister to Euploca (formerly Hilgeria, Schleidenia, Heliotropium sect. Orthostachys). The remaining sections of Heliotropium, Tournefortia sect. Tournefortia and the three small genera Argusia, Ceballosia, and Nogalia, segregated from Heliotropium, constitute the second large clade. Argusia, Ceballosia, and Nogalia cluster within clades of Heliotropium and therefore are reincluded in this genus. Within Heliotropium, the species of former Tournefortia sect. Tournefortia represent a lineage of tropical New World Heliotropium species, growing in humid environments, whereas all other Heliotropium species are found in semi-arid habitats. Before new combinations in the genus Heliotropium are made for "Tournefortia", the exact relationship within New World Heliotropium needs to be resolved, and a revision of "Tournefortia" is inevitable. We advocate maintaining the genus Tournefortia, which is easily defined, and we conclude that under this definition the genus Heliotropium is paraphyletic. Five genera are thus accepted; 22 new combinations within Heliotropiaceae are presented.

1 aHilger, HH1 aDiane, N uhttps://boraginales.myspecies.info/node/6001067nas a2200217 4500008004100000245010000041210006900141300001400210490000700224520042800231653001000659653001700669653001700686653001100703653001800714653001500732653002500747100001500772700001500787856004700802 2000 eng d00aPontechium: a new genus distinct from Echium and Lobostemon (Boraginaceae)0 aiPontechiumi a new genus distinct from iEchiumi and iLobostemoni a737–7460 v493 a

Plastid (trnL(UAA) intron, trnL(UAA)-trnF(GAA) spacer) and nuclear (ITS1) DNA sequence analyses do not only confirm the separate position of „Echium maculatum L." which this species holds within the genus Echium due to the presence of a bicapitate rather than a bifid stigma, but necessitate the exclusion of the taxon from the genus Echium (Boraginaceae) and the description of a new genus, Pontechium.

10a{DNA}10aBoraginaceae10aEchiostachys10aEchium10aLithospermeae10aLobostemon10aPontechium gen. nov.1 aHilger, HH1 aBöhle, UR uhttps://boraginales.myspecies.info/node/61