02636nas a2200193 4500008004100000245023100041210006900272490000800341520190000349653001702249653002002266653001302286653002402299653001602323653002602339100001502365700002002380856004202400 2009 eng d00aPhylogenetic relationships of the monotypic genera Halacsya and Paramoltkia and the origins of serpentine adaptation in circum- mediterranean Lithospermeae (Boraginaceae): insights from ITS and matK DNA sequences0 aPhylogenetic relationships of the monotypic genera iHalacsyai an0 v58 3 a

Halacsya and Paramoltkia are monotypic and partially sympatric genera in the Balkans, with no clear relation-
ships among extant Lithospermeae due to striking morphological autapomorphies and scarcity of phylogenetic
analyses in this group. The two species H. sendtneri and P. doerfleri show a strict selectivity for serpentine soils,
posing the question whether this edaphic specialization reflects a common ancestry or a parallel process of adap-
tive evolution in unrelated lineages. DNA sequences from the nuclear ITS and chloroplast matK regions were
generated from multiple accessions of Halacsya and Paramoltkia, and from representatives of 16 other genera
of Mediterranean Lithospermeae. SEM analyses of pollen morphology were also conducted to test relationships
indicated by molecular phylogenies. Parsimony analyses retrieved a clade of morphologically well differentiated
monotypic Lithospermeae including Halacsya and Paramoltkia. ITS provided a better resolution of relationships
and showed the two genera to be sistergroups close to Mairetis and Moltkiopsis, and no affinity to Moltkia as sup-
posed by past authors. Pollen characters corroborated the phylogenetic link between the two Balkan genera. Five
further monophyletic clades were recognised: Onosma-Echium, Moltkia, Lithospermum s.l., Arnebia-Macrotomia,
and Alkanna-Podonosma. Mapping the edaphic preferences of Lithospermeae onto molecular cladograms showed
that serpentinophytism as an obligate condition originated separately in the clade of monotypic genera and in
that of Onosma-Echium. In Halacsya and Paramoltkia it represents an early ecological synapomorphy which
probably originated in situ from non-serpentine ancestors related to Moltkiopsis and Mairetis.

 

10aBoraginaceae10aedaphic habitat10aHalacsya10amolecular phylogeny10aParamoltkia10aserpentine adaptation1 aCecchi, L.1 aSelvi, Federico u http://www.jstor.org/stable/2775693901875nas a2200193 4500008004100000245007700041210006900118490000800187520130900195653001701504653001601521653001801537653001401555653001601569653001301585100002001598700001601618856004701634 2012 eng d00aA synopsis of the genus Cynoglossum (Boraginaceae-Cynoglosseae) in Italy0 asynopsis of the genus Cynoglossum BoraginaceaeCynoglosseae in It0 v1463 a

Based on field and herbarium investigations, a systematic synthesis of the critical genus Cynoglossum (Boraginaceae) in Italy is
provided. Types, diagnostic characters and original iconographies are given for each native species, together with a revised
analytical key. Regional distributions are outlined and a list of selected vouchers is included. Nine species are native to the
Italian territory: Cynoglossum barbaricinum, Cynoglossum cheirifolium, Cynoglossum clandestinum, C. columnae, C. creticum, C.
magellense, C. montanum, C. nebrodense and C. officinale. A new subspecies, named C. nebrodense ssp. lucanum, is described
from some calcareous massifs in S Basilicata and N Calabria. This taxon is morphologically and geographically distinct from
the typical C. nebrodense s.str. In addition, C. dioscoridis and C. germanicum, often reported from Italy, cannot be confirmed
and are apparently to be excluded from the Italian flora. Old herbarium records of C. amabile Stapf & Drummond and C.
coelestinum Lindb. [: Paracaryopsis coelestina (Lindl.) R.R. Mill] from northeast Italy are reported for the first time, although
the presence of these two exotic species in Italy was probably occasional.
 

10aBoraginaceae10aCynoglossum10aItalian flora10akaryology10asystematics10aTaxonomy1 aSelvi, Federico1 aSutorĂ½, K. uhttps://boraginales.myspecies.info/node/3302329nas a2200205 4500008004100000245007800041210006900119490000700188520173000195653001701925653001301942653002501955653002401980653001602004653001302020100002002033700001502053700001402068856004102082 2009 eng d00aPhylogeny, karyotype evolution and taxonomy of Cerinthe L. (Boraginaceae)0 aPhylogeny karyotype evolution and taxonomy of Cerinthe L Boragin0 v583 a

A phylogenetic and karyological analysis of the small and poorly known genus Cerinthe L. (Boraginaceae-
Lithospermeae) was performed using ITS sequences and standard chromosomal techniques. All taxa are
diploid with 2n = 16 or 2n = 18 and show a variable degree of infraspecific variation, in particular in the
polymorphic C. major and C. minor. Change in base number is associated with an early split between the
two well-supported clades of C. major, corresponding to Cerinthe sect. Cerinthe, and that of all other taxa
belonging to C. sect. Ceranthe, with the base x = 8 found only in the strictly annual C. major group, and
x = 9 in the other five species of the genus: C. minor, C. glabra, C. tenuiflora, C. retorta and C. palaestina.
The latter section is subdivided into the E Mediterranean, annual lineage of C. palaestina–C. retorta and
the mainly continental, perennial group of C. minor–C. glabra, the sister of which is the Corsican endemic
C. tenuiflora. The hypothesis that x = 9 represents the primary haploid number and x = 8 is derived through
descending aneuploidy, is discussed. A taxonomic revision of the genus is provided and the following formal
taxonomic changes are proposed: C. major L. subsp. oranensis (Batt.) Selvi & Cecchi, stat. nov.; C. major
L. subsp. purpurascens (Boiss.) Selvi & Cecchi, stat. nov.; C. minor L. subsp. cleiostoma (Boiss. & Sprun.)
Selvi & Cecchi, stat. nov. Cerinthe tenuiflora, C. retorta and the poorly known C. palaestina are well-defined
species with little internal variation.

 

10aBoraginaceae10aCerinthe10achromosome evolution10amolecular phylogeny10asystematics10aTaxonomy1 aSelvi, Federico1 aCecchi, L.1 aCoppi, A. uhttp://www.jstor.org/stable/27757019