02094nas a2200181 4500008004100000245009900041210007100140300001400211490000800225520146600233653006301699100002201762700002101784700001501805700002801820700001601848856004801864 2016 eng d00aTowards a monophyletic Omphalodes—or an expansion of North American Mimophytum0 aTowards a monophyletic iOmphalodesi—or an expansion of North Ame a131–1440 v2883 a

Molecular studies have recently led to major changes to the systematics of Boraginaceae and changed our perception of fruit morphology, previously used as the most important character in Boraginaceae classification. Navel-shaped nutlets as the key character for Omphalodes turned out to be highly homoplasious, thus the genus was revealed to be polyphyletic. After the recent reclassifications, the genus Omphalodes is still left paraphyletic, since the remaining taxa correspond to two different lineages: a clade of essentially Mediterranean perennials including the type species (O. verna) and a clade with the annual to subperennial North American taxa including the genus Mimophytum . We here advocate transferring all taxa of the North American clade into the genus Mimophytum, subdividing the genus Omphalodes into morphologically coherent and monophyletic entities. Mimophytum was initially removed from Omphalodes based on its glochidiate nutlets, but this character state does not define a monophyletic entity. In overall morphology, the redefined genus Mimophytum is highly homogeneous and easily differentiated from the other clades: it has a lax growth habit, and leaves are long-petiolate, abruptly contracted at the base into a more or less cordate base, and with a fine pubescence of erect trichomes. After the proposed rearrangements, only O. erecta remains as a problematic species whose systematic placement is still to be found out.

10aEudicots, Boraginaceae, navel seeds, Sierra Madre Oriental1 aHolstein, Norbert1 aChacón, Juliana1 aOtero, Ana1 aJiménez-Mejías, Pedro1 aWeigend, M. uhttps://boraginales.myspecies.info/node/12101393nas a2200241 4500008004100000245009900041210006900140260001500209300000700224490000800231520068700239653001300926653001300939653001300952653001000965653002500975653001801000653001701018653001401035100002201049700001601071856006401087 2016 eng d00aLectotypifications and nomenclatural notes in the Boraginaceae s.str. for the Flora of Ecuador0 aLectotypifications and nomenclatural notes in the Boraginaceae s c05/10/2016 a730 v2783 a

During the preparation of a taxonomic treatment concerning Boraginaceae s.str. for the Flora of Ecuador, we found that there are eleven species currently known in that area. We herewith designate lectotypes for some of the names concerned: Cynoglossum revolutum and Cynoglossum mexicanum (both Hackelia revoluta), Amsinckia hispida and Eritrichium mandonii (both Amsinckia calycina), and Anchusa linifolia and Lithospermum alpinum (both Plagiobothrys linifolius). The complex history of the type material of various synonyms of Plagiobothrys linifolius based on material by Bonpland and Humboldt is briefly discussed.

10aBonpland10aEudicots10aHumboldt10aKunth10aRuiz López & Pavón10aSouth America10atypification10aWilldenow1 aHolstein, Norbert1 aWeigend, M. uhttp://biotaxa.org/Phytotaxa/article/view/phytotaxa.278.1.902602nas a2200241 4500008004100000245017500041210007100216250001600287260003200303300001400335490000800349520178600357653002102143653000802164653002402172653001302196653002002209100002202229700002102251700002402272700001602296856004802312 2016 en d00aNo longer shipwrecked—Selkirkia (Boraginaceae) back on the mainland with generic rearrangements in South American “Omphalodes” based on molecular data0 aNo longer shipwrecked—iSelkirkiai Boraginaceae back on the mainl a24 Aug 2016 bMagnolia Pressc24 Aug 2016 a231–2510 v2703 a

The genus Omphalodes (Boraginaceae) has recently been shown to be polyphyletic. Two distantly related lineages have already been segregated into the genera Memoremea (Central Europe) and Nihon (East Asia), respectively. We expanded the taxon sampling in the Omphalodeae and confirm that the genus is still paraphyletic to the two monotypic genera Selkirkia from the Juan Fernández Islands off the coast of Chile and Myosotidium from Chatham Island off the coast of New Zealand, plus two South American species currently assigned to Cynoglossum, and one species recently segregated from the latter genus as Mapuchea. Four clades are retrieved in a narrowly delimited Omphalodes group: 1) Iberodes, the annual southwestern European species of Omphalodes s.l. that have been recently segregated into this genus, 2) Omphalodes s.str., perennial western Eurasian species (including the type species of the genus), 3) the North American species of Omphalodes, and 4) the southern hemispheric Myosotidium as sister to a monophyletic group with Mapuchea plus the two other South American species of “Cynoglossum” and the island shrub Selkirkia berteroi. We argue that the taxa of this latter clade are best placed into an expanded genus Selkirkia. Selkirkia then represents a morphologically coherent entity with glochidiate nutlets. Its considerable difference in vegetative morphology to Myosotidium is easily explained by the highly divergent habitats the respective plants occupy. Lectotypifications, illustrations, and descriptions are provided for Myosotidium and the species of the expanded genus Selkirkia.

10aisland woodiness10aITS10amolecular phylogeny10aTaxonomy10atrnL–trnL–F1 aHolstein, Norbert1 aChacón, Juliana1 aHilger, Hartmut, H.1 aWeigend, M. uhttps://boraginales.myspecies.info/node/11600581nas a2200181 4500008004100000245007800041210006900119260001200188300001600200490000700216653002200223653002100245653002200266653002500288100002200313700001600335856004800351 2015 eng d00a(2388) Proposal to reject the name Myosotis collina (Boraginaceae)0 a2388 Proposal to reject the name iMyosotis collinai Boraginaceae c10/2015 a1058–10590 v6410aMyosotis arvensis10aMyosotis collina10aMyosotis discolor10aMyosotis ramosissima1 aHolstein, Norbert1 aWeigend, M. uhttps://boraginales.myspecies.info/node/112